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@gbj_tester
About 10% more aerodynamic drag, seemingly entirely due to the fat aluminium spokes. If you factor that into a typical race speed/gradient profile, along with the much lower weight of the Ksyrium, it turns out that it takes more energy to ride the Ksyriums at a given speed than the Aksiums, unless you're riding a hill climb where the lighter weight wheel gains an advantage.
Aksiums are hardly a paragon of the wheelbuilders art, but since they are Mavic's cheapest wheel and also the one they gave to owners of the (even slower) exploding R-Sys, the comparison is enlightening.