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Guigs415
I've got 28's on my 585^ but that's as big as will fit.
In my experience the older GP4000's blew up bigger than the current GP5000's, and though I've not got a 481 I'm sure you'd fit 25c GP5000 no problem if your GP4000's fit, you might even manage a 28c GP5000 in there, but I'd wait for confirmation from a 481 owner.