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Just clarifying on spanking in case you're actually hitting them. You can pick up brightbloom seeds and throw them (hold down R, hit up, select object to throw, release up, then release R when you're aimed).
Had no idea about the water matching up with the 'impassable' bits below. ' = because I've actually found tiny navigable tunnels through several of the bigger ones. I've got all the 120 lightroots but still only 102 shrines.
CYOA
NotThamesWater
There are bosses in this game?
I'm still hitting up every lightroot in the depths. With 3 hearts. I don't know why.
When I had the revelation that the Lightfoot's and temples are at the same coordinates, it did become a bit easier, though, and I didn't have to spank every single bright bulb any more.
As did realising that water in the main level translated to impassable cliffs down below.