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Stonehedge-
There's not much difference between the DM and MailOnline - they're both edited by Paul Dacre. They're separate so that the DM can claim the moral high ground without appearing to be hypocritical while MailOnline does sex'n'tits'n'arses. MoS is edited by Geordie Greig, and as mentioned above, he and 'double-cunting' Dacre do not get along. At all.
Brommers
The Mail on Sunday has a different editorial management team to the Daily Mail. Which shows quite a lot.