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Those are all 255, apart from Fat12
And if you've got a 16MB file size limit then dropping from 255 to 254 character filenames is the least of your worries :DThere's also the MAX_PATH limit that complicates things sometimes
https://msdn.microsoft.com/en-gb/library/windows/desktop/aa365247(v=vs.85).aspx#maxpath
Depends on filesystem:
http://www.ntfs.com/ntfs_vs_fat.htm