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@alibi
I think you are right to question that.
The sentence quoted is poorly worded. I think they are three separate statements, they have flipped the tube to 'soften the ride', also because they 'mate better' and 'look better'. I don't think they are saying as they mate better therefore a softer ride. Stiffer if anything, surely?
If you read up a little in the article they mention more on the topic.
My understanding, the oval tubes are stiffer across the longer section. If the flatter section is at the headtube there should be slightly more flex come through from the fork/head tube to soften the ride.