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@ewanmac
^ largely this. The front of the saddle needs to be (?)3cm behind the bottom bracket to be UCI legal whereas triathletes have no such rule and therefore tend to have their saddles further forward to enable a slightly different engagement of muscles to save their legs for the run. Also, all of the "add ons" on the Tri bike (i.e. behind saddle box and maybe even the toptube bento box). are also non UCI compliant - they are designed primarily for long distance triathletes (Ironman etc) who need to carry food and spares with them while racing.
FWIW, I believe that the "Tri bike" is the latest incarnation of the SC range and was only released recently, while the leopard "TT bike" is the older model. The new model allows even more Tri specific/non-UCI features such as a fin that reaches around below the BB and shields the rear brake. Also, I believe that strictly speaking Trek always designed the SC range primarily as Tri bikes that could be used for TTs as distinct to most brands that primarily build TT bikes that people then use for Tri.
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