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@Sanddancer
A while back I was doing my club's slower, Saturday, rides as well as the proper Sunday ones. I enjoyed it for the craic, but the rides were a bit dull on my road bikes so I started doing them on my 464 to make it more interesting. ~65-75 miles every Saturday over crap, rutted, pot-holed Northumberland roads and never had an issue with comfort. That's built with 28 spoke high flange hubs to H+Son SL42s, Deda Pista bars and Thomson post; not exactly components that would lessen the harshness of the frame.
Edit: I do run 25mm Maxxis Refuses, mind.