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@Rik_Van_Looy
The way it was always explained to me was that when you're going quick on the flat a steep effective seattube angle helps to get your weight onto the pedals and gain power on the downstroke - hence the popularity of steep seat angles and inverted seatposts in triathalon.
When you're spinning up long climbs you want to be sat well back comfortably turning nice clean circles.
Hence TT & Crit bikes tended to be much steeper than "normal" road race geometry