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@Smallfurry
I'm not 100%. But I guess its down to cable pull. 9spd shimano works great when you mix and match. This made it great for trekking/touring/commuting etc.
But when the MTB stuff went 10spd they went with what they call Dyna-sys.
http://bike.shimano.com/publish/content/global_cycle/en/us/index/products/dynasys.html
At the same time 10spd Sram stuff is all compatible now with 'exact actuation'. I have a TT lever, and a X0 rear mech I'm going to be running together.