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@Vince
I'm pretty sure it's all 653 tubing, including forks - I've interrogated Google and all other 653 Cougars I have spotted are all 653, like this one:
http://www.ebay.co.uk/itm/cougar-reynolds-653-road-frame-fork-54cm-fixie-campag-/130540296858
Cheers
Vince