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@nneil
I contacted Chain Reaction cycles, who sold me the tyres. They said that the Schwalbe Snow Stud is particularly troublesome to fit (I must say that they have been very helpful in general). Of course Schwalbe think the tyre should fit fine just fine.
Having played with the two tyres on the rim it is clear that the Specialized one hooks on readily and firmly. The Schwalbe doesn't: the wire "hook" built into the bead just isn't that pronounced.
I have ordered the WTB rim. I note it has an "International Bead Seat", which is supposed to help hold the tyre on the rim. We shall see.