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Only data point I can offer is getting train back from Moreton-in-Marsh, tiddly station in Cotswolds last year after a solstice ride.
Travelled super off peak at like mid-day on a Weds. Had carbon wheels so wasn't mega enthused about hanging bike up. Guard decided to make a real song and dance about it on the largely empty train, insisted on making me hang it up despite someone's wide-barred MTB already taking up 90% of the space in the little bike cupboard.
Friends on another train behind said no issues like this whatsoever.
tl;dr: your life is in the hands of the particular guard on the train you're on. If you're in no rush, plead ignorance and wait for the one behind?
o rly? i know their racks are shit but if i reserve a space and cite ignorance? “oh i didnt know they were vertical racks” .. journey is off peak, only 1.5hrs