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I did a bit of testing, and I think I was wrong when I said uploads aren't changed - something is definitely modifying (downscaling and EXIF stripping) images uploaded to LFGSS.
At first I thought it was Chrome just stripping EXIF data for privacy reasons, (which apparently is the default now, TIL), but then I uploaded a photo with curl and it was still stripped and down-scaled. This must be a new change because it wasn't always so.
My Flickr account is dormant at the moment so I can’t actually upload anything to it without coughing-up.
Could it be to do with the colourspace being P3 and being converted by microcosm? I could try uploading different colourspace versions of the same image in a different thread and seeing if that makes a difference.