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They are a real mix of old nails, newer nails, screws and even no nails.
I’m moving around a lot of the boards to better stagger how they are because at the moment there are a lot of ugly clear runs of cuts and joins.
Will re-nail down the boards. Where there are any old nail holes that go un-used is the best thing to just mix up some sawdust and pva and fill with that?
Or bang them all the way through the board down in to the joist with a punch if you can, then the board will lift off and there'll be a small hole left. Assuming they are old flooring nails, not modern ones with a big round head
EDIT - zoomed in to the photos a bit more, I'd just bang them in flat