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It might not change the feel but if you're putting a measureably faster tyre on then it will reduce your rolling resistance so you will go faster for same energy. If you only use one fast tyre, sure you only get half the benefit, but it's still a benefit and if your aim is to ride fast you might need that front grip still.
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I wouldn't have noticed the Crust's wheels at all from just the photo.
Wheels and tyres are the same size in the photo. @kjlem was talking about the stated tyre clearance.
It is odd. The only thing I can think of is that they wanted a tucked rear end, and the front is a result of the fork crown and legs choice. But I don’t know enough about frame building to prove that.
Also lugs. Doesn’t that limit your geo freedom?
Because I'm a unique fucking snowflake, I have a reverse mullet - 45mm front and 50mm rear.