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During the winter the upvc windows are generally closed, or open on the locked but just open position so I’m pretty sure it’s not that.
From what the previous owner said, chimneys are still open/usable. Occasionally little bits fall down past the insulation board blocking the draft downstairs.
It could be floor voids, but it does seem like it’s emanating from those specific rooms and the fireplace, although I may be imagining it.
cjr
If the smell isn’t coming in through the windows then it might be through voids in the floor and ceiling.
Are the chimneys still present and are they capped?