one or more of the cups/cones out of alignment meaning the space for the balls widens or narrows depending on where the bars are pointing.
Cheers - is there anything else that might cause this other than cross-threading?
That's a strong candidate, as the treads would be keeping it mostly in line. That or one of the cups have shifted from the time it was cross threaded, and putting the nut/cone back on better aligned now means they are not parallel.
@youClown started
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one or more of the cups/cones out of alignment meaning the space for the balls widens or narrows depending on where the bars are pointing.