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It looked like there was one there at some point, but had been boarded over.
One of the previous owners seems to have been a little obsessed stopping drafts. The old in-room vents were stuffed with cotton wool and painted over. Every gap around the floor and skirting is full of silicon, there's bits of expanding foam all over the place. A wall vent in the kitchen has a sheet of paper stuck over it and has then been painted over (this is not a papered wall). The list goes on...
atk
Very useful, thanks. I will assume it originally vented at the step down.
My preference is to break out the concrete and replace with a new insulated raised timber floor but I imagine that would be quite fiddly and expensive.