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Except there aren't the same rapid traction changes you get with those surfaces. I don't know about your turbo trainers but none of mine respond so rapidly that I have a jerky 'off road' pedal stroke.
Riding up that Alp du Zwift thing definitely feels different to the real world but all the other routes where the gradient changes aren't so frequent feel pretty normal, especially if you keep your cadence up when the gradient changes.
hippy
I would say that most people are forced to pedal differently. It's the same difference between peddling up a smooth flat road vs up a steep hill or through sand/mud - you can be doing the same watts and rpm but the pedal stroke and muscle demands are different.