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Here's my question: is this an oak (or other wood) board or just dodgy-looking pine?
To me looks like somebody replaced that board with some pine and sanded it level when redoing the floors. The wood filler in the adjacent board is probably part of the story.
the coloration could be a result of anything from simple time to somebody trying to stain it to match very poorly.
The rest of your boards looks like heart pine to me.
I am not a hardwood floor expert, but I have very very old pine and redwood boards in my house and I've been through this with several experts.
Just a guess.
Here's my question: is this an oak (or other wood) board or just dodgy-looking pine?