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Managed to clear 42 x 700C (WTB resolute) without mudguards. The limiting factor isn't actually the clearance to the stays, but the clearance between the front derailleur lever and the tyre when on high gear (clearance is at best 2-3 mm)...
With mudguards, I run 38 x 700C Gravelkings SK without any problems.
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Just curious, what's the biggest tyre anyone has fit on their Croix de Fer?