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Yeah, can be a pain. I think there are one or two direct trains to inverness from London during the day too.
The route is now published on their site. Not sure if he is going to run as a mandatory route like the 400 in the organisers series.
https://ridewithgps.com/routes/28688630
If you still looking for a Scottish 600. There is another running from Inverness on 8 June. Pretty tough route.
http://www.aukweb.net/events/detail/19-953/
Sleeper train options to/from Inverness from London.