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Cheers for the feedback (you too @dancing james).
The rim in question is a Stan's ZTR Grail which is BST-R, which is probably at odds with the IRC Formula Pro X-guard in question (though I haven't died a jizzy death yet).
Happily I don't have to burden you with the old question of 'why does my tyre deflate', as these seem to hold the air considerably better than Schwalbe nightmares.
You may be missing the bead retaining lip (arrowed in the drawing). It's in the UST standard, but seems to have gone astray in a great many so-called "tubeless ready" rims.