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the UST ones use FORE cups
You can get UST certification without having the undrilled bed, as long as you pay the fee and pass the dimensional tolerance tests. I'm not even sure that you have to pass an air seal test, UST is mostly about making sure every UST tyre will mount on, and stay on, every UST rim of appropriate width.
Sure? I am looking at google results of 421 and found this (scroll down):
Now I will need to call them up and confirm if they are the same rims ..
Also this:
Then these two contradictory results ..
https://www.bike-components.de/en/Mavic/EN-421-Disc-29-Felge-Modell-2016-Auslaufmodell-p45720/
http://www.chainreactioncycles.com/mavic-en421-disc-mtb-rim-2015/rp-prod127198
Mavic are such cunts for not documenting things properly.