Citation needed. The rolling resistance is very close, and the tread on the GP4000S seems to act as a trip, so some wheels have less drag with a 4000 than they do with a completely slick tyre.
Not off the top of my head, and I'm not about to embark on a lengthy quest on my phone, but that new Swiss wheel company (Swissside?) or maybe Flo tested different tyres and one or both found the 4000s had good aeros IIRC.
Citation needed. The rolling resistance is very close, and the tread on the GP4000S seems to act as a trip, so some wheels have less drag with a 4000 than they do with a completely slick tyre.