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Fair enough, the backward facing footage didn't really show anything conclusive (as I said at the time).
Still, two riders not really looking where they are going and ending up swerving towards a motorbike that was swerving towards them.
Footage here: http://bleacherreport.com/articles/2529773-jakob-fuglsang-finishes-stage-18-in-5th-place-after-being-hit-by-motorcycle
Either hit a rut in the road or, possibly, he swerved way out and clipped the back of motorbike that just went past. Can't tell from the limited footage there was.
Either way it was his fault.
It was Fuglsang.