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@edscoble
seat tube angle change the reach, the slacker seat tube add at least a centimetre to the length.
V4 have 72.5 degree ST, with 561cm TT.
V3 have 73.5 degree ST, with 551cm TT.
V3 have 1 degree steeper seat tube, with 1cm shorter reach.
However one could argue that it always been the V4 sizes in the predecessor as On-one isn't the brightest when it come to publishing their geometry (like Dolan).